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FAMILY Cricetidae

Author:Fischer, 1817.
Citation:Mém. Soc. Imp. Nat. Moscow, 5: 372.
Comments:

The cricetid-murid question has persisted as the dominant theme, or uncertainty, involving the higher level classification of muroid rodents over the past century (see summaries in Carleton, 1980, and Carleton and Musser, 1984): in essence, should the various subfamilies be approximately equally apportioned between Cricetidae and Muridae (Miller and Gidley, 1918; Simpson, 1945) or should most be placed under an all encompassing Muridae (Alston, 1876; Ellerman, 1940; Thomas, 1896—even these studies, however, accorded the fossorially specialized forms separate familial rank; i.e., the spalacines, rhizomyines, and myospalacines in various combinations). While the inclusive view of Muridae has gained acceptance over the latter half of the 20th century (Hershkovitz, 1962; Hooper and Musser, 1964a; McKenna and Bell, 1997), there are notable departures (Chaline et al., 1977; Lavocat, 1978), and Reig (1980, 1981, 1984), in particular, has steadfastly espoused recognition of a separate Cricetidae, the contents of which correspond in large part to the taxa covered here. Principal subfamilial exceptions to Reig’s coverage are exclusion of certain African clades (Cricetomyinae, Dendromurinae, Nesomyinae), here arranged in a broadly defined Nesomyidae (as per Lavocat, 1973, 1978), and inclusion of Arvicolinae and Lophiomyinae, each of which Reig regarded as separate families. Accepting those adjustments, monophyly of the family finds general support in phylogenetic evaluation of genetic sequence data (Dubois et al., 1999; Michaux and Catzeflis, 2000; Michaux et al., 2001b; Robinson et al., 1997). Nevertheless, this cladistic hypothesis of Cricetidae should be viewed as provisional, and taxonomic sampling must be considerably expanded to reinforce its validity, notably with addition of groups such as Lophiomyinae and many more cladistically older members drawn from Cricetinae, Sigmodontinae, and Tylomyinae.

Although monophyletic on molecular trees, unambiguous diagnosis of so large and heterogeneous an assemblage would prove challenging with the morphological character base currently referenced in phylogenetic studies. We note that all members possess a biserial arrangement of the molar cusps, with retention of a longitudinal connection (mure/murid) among them and formation of a discrete anterocone(id) on the first molars. Such character states are likely ancestral for the superfamily, as are other traits exhibited by many cricetid species, in some cases a majority of them (three-rooted upper and two-rooted lower molars; possession of mesolophs(ids); malleus configuration parallel; tegmen tympani conformation; complete carotid circulatory pattern; entepicondylar foramen present; stomach unilocular-hemiglandular; full complement of male reproductive glands). However, parallel evolution of derived conditions for each of these characters is apparent at lower taxonomic levels within Cricetidae (subfamilies and tribes) and among other families of Muroidea as currently understood.

The fossil record of "cricetids" is comparatively rich in taxa, deep in time, and broad in geographic representation, and their phylogenetic systematics is correspondingly complicated. Several groups—e.g., cricetodontines, eucricetodontines, paracricetodontines, and cricetopines—have been variously acknowledged as families separate from Cricetidae; McKenna and Bell (1997) recapitulated family-group rankings used for such extinct cricetids and retained most as tribes or subfamilies within Muridae sensu lato. Also see overview by Carleton and Musser (1984) and regional treatments by Freudenthal et al. (1992), Hugueney (1999), Kälin (1999), Korth (1994), Lavocat (1978), Martin (1980), Mein and Freudenthal (1971), and Rümmel (1999). The evolutionary fate of many of these extinct groups as progenitor to living cricetid assemblages is generally unclear and-or much disputed; where some phyletic link is plausibly documented, usually not until Miocene in age, we mention those groups in the subfamily comments. Geological range from the early Oligocene to Recent of Europe, late Eocene to Recent of Asia, late Miocene to Recent of Africa, late Eocene to Recent of North America, and early Pliocene to Recent of South America (McKenna and Bell, 1997).

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Offspring:

SUBFAMILY Arvicolinae

GENUS Alticola

SUBGENUS Alticola

SPECIES albicaudus

SPECIES argentatus

SPECIES barakshin

SPECIES montosa

SPECIES roylei

SPECIES semicanus

SPECIES stoliczkanus

SPECIES tuvinicus

SUBGENUS Aschizomys

SPECIES lemminus

SPECIES macrotis

SPECIES olchonensis

SUBGENUS Platycranius

SPECIES strelzowi

GENUS Arborimus

SPECIES albipes

SPECIES longicaudus

SPECIES pomo

GENUS Arvicola

SPECIES amphibius

SPECIES sapidus

SPECIES scherman

GENUS Blanfordimys

SPECIES afghanus

SPECIES bucharensis

GENUS Caryomys

SPECIES eva

SPECIES inez

GENUS Chionomys

SPECIES gud

SPECIES nivalis

SPECIES roberti

GENUS Dicrostonyx

SPECIES groenlandicus

SPECIES hudsonius

SPECIES nelsoni

SPECIES nunatakensis

SPECIES richardsoni

SPECIES torquatus

SPECIES unalascensis

SPECIES vinogradovi

GENUS Dinaromys

SPECIES bogdanovi

GENUS Ellobius

SUBGENUS Ellobius

SPECIES alaicus

SPECIES talpinus

SPECIES tancrei

SUBGENUS Afganomys

SPECIES fuscocapillus

SPECIES lutescens

GENUS Eolagurus

SPECIES luteus

SPECIES przewalskii

GENUS Eothenomys

SPECIES cachinus

SPECIES chinensis

SPECIES custos

SPECIES melanogaster

SPECIES miletus

SPECIES olitor

SPECIES proditor

SPECIES wardi

GENUS Hyperacrius

SPECIES fertilis

SPECIES wynnei

GENUS Lagurus

SPECIES lagurus

GENUS Lasiopodomys

SPECIES brandtii

SPECIES fuscus

SPECIES mandarinus

GENUS Lemmiscus

SPECIES curtatus

GENUS Lemmus

SPECIES amurensis

SPECIES lemmus

SPECIES portenkoi

SPECIES sibiricus

SPECIES trimucronatus

GENUS Microtus

SPECIES abbreviatus

SPECIES californicus

SPECIES chrotorrhinus

SPECIES guatemalensis

SPECIES longicaudus

SPECIES mexicanus

SPECIES miurus

SPECIES richardsoni

SPECIES umbrosus

SPECIES xanthognathus

SUBGENUS Microtus

SPECIES agrestis

SPECIES anatolicus

SPECIES arvalis

SPECIES cabrerae

SPECIES dogramacii

SPECIES guentheri

SPECIES ilaeus

SPECIES irani

SPECIES levis

SPECIES paradoxus

SPECIES qazvinensis

SPECIES schidlovskii

SPECIES socialis

SPECIES tatricus

SPECIES transcaspicus

SUBGENUS Terricola

SPECIES bavaricus

SPECIES brachycercus

SPECIES daghestanicus

SPECIES duodecimcostatus

SPECIES felteni

SPECIES gerbei

SPECIES liechtensteini

SPECIES lusitanicus

SPECIES majori

SPECIES multiplex

SPECIES savii

SPECIES schelkovnikovi

SPECIES subterraneus

SPECIES thomasi

SUBGENUS Mynomes

SPECIES breweri

SPECIES canicaudus

SPECIES montanus

SPECIES oregoni

SPECIES pennsylvanicus

SPECIES townsendii

SUBGENUS Alexandromys

SPECIES clarkei

SPECIES evoronensis

SPECIES fortis

SPECIES kikuchii

SPECIES limnophilus

SPECIES maximowiczii

SPECIES middendorffii

SPECIES mongolicus

SPECIES montebelli

SPECIES mujanensis

SPECIES oeconomus

SPECIES sachalinensis

SUBGENUS Stenocranius

SPECIES gregalis

SUBGENUS Pitymys

SPECIES oaxacensis

SPECIES pinetorum

SPECIES quasiater

SUBGENUS Pedomys

SPECIES ochrogaster

GENUS Myodes

SPECIES andersoni

SPECIES californicus

SPECIES centralis

SPECIES gapperi

SPECIES glareolus

SPECIES imaizumii

SPECIES regulus

SPECIES rex

SPECIES rufocanus

SPECIES rutilus

SPECIES shanseius

SPECIES smithii

GENUS Myopus

SPECIES schisticolor

GENUS Neodon

SPECIES forresti

SPECIES irene

SPECIES juldaschi

SPECIES sikimensis

GENUS Neofiber

SPECIES alleni

GENUS Ondatra

SPECIES zibethicus

GENUS Phaiomys

SPECIES leucurus

GENUS Phenacomys

SPECIES intermedius

SPECIES ungava

GENUS Proedromys

SPECIES bedfordi

GENUS Prometheomys

SPECIES schaposchnikowi

GENUS Synaptomys

SUBGENUS Synaptomys

SPECIES cooperi

SUBGENUS Mictomys

SPECIES borealis

GENUS Volemys

SPECIES millicens

SPECIES musseri

SUBFAMILY Cricetinae

GENUS Allocricetulus

SPECIES curtatus

SPECIES eversmanni

GENUS Cansumys

SPECIES canus

GENUS Cricetulus

SPECIES alticola

SPECIES barabensis

SPECIES kamensis

SPECIES longicaudatus

SPECIES migratorius

SPECIES sokolovi

GENUS Cricetus

SPECIES cricetus

GENUS Mesocricetus

SPECIES auratus

SPECIES brandti

SPECIES newtoni

SPECIES raddei

GENUS Phodopus

SPECIES campbelli

SPECIES roborovskii

SPECIES sungorus

GENUS Tscherskia

SPECIES triton

SUBFAMILY Lophiomyinae

GENUS Lophiomys

SPECIES imhausi

SUBFAMILY Neotominae

GENUS Baiomys

SPECIES musculus

SPECIES taylori

GENUS Habromys

SPECIES chinanteco

SPECIES delicatulus

SPECIES ixtlani

SPECIES lepturus

SPECIES lophurus

SPECIES simulatus

GENUS Hodomys

SPECIES alleni

GENUS Isthmomys

SPECIES flavidus

SPECIES pirrensis

GENUS Megadontomys

SPECIES cryophilus

SPECIES nelsoni

SPECIES thomasi

GENUS Nelsonia

SPECIES goldmani

SPECIES neotomodon

GENUS Neotoma

SUBGENUS Neotoma

SPECIES albigula

SPECIES angustapalata

SPECIES anthonyi

SPECIES bryanti

SPECIES bunkeri

SPECIES chrysomelas

SPECIES devia

SPECIES floridana

SPECIES fuscipes

SPECIES goldmani

SPECIES lepida

SPECIES leucodon

SPECIES macrotis

SPECIES magister

SPECIES martinensis

SPECIES mexicana

SPECIES micropus

SPECIES nelsoni

SPECIES palatina

SPECIES stephensi

SUBGENUS Teonoma

SPECIES cinerea

SUBGENUS Teanopus

SPECIES phenax

GENUS Neotomodon

SPECIES alstoni

GENUS Ochrotomys

SPECIES nuttalli

GENUS Onychomys

SPECIES arenicola

SPECIES leucogaster

SPECIES torridus

GENUS Osgoodomys

SPECIES banderanus

GENUS Peromyscus

SPECIES attwateri

SPECIES aztecus

SPECIES beatae

SPECIES boylii

SPECIES bullatus

SPECIES californicus

SPECIES caniceps

SPECIES crinitus

SPECIES dickeyi

SPECIES difficilis

SPECIES eremicus

SPECIES eva

SPECIES fraterculus

SPECIES furvus

SPECIES gossypinus

SPECIES grandis

SPECIES gratus

SPECIES guardia

SPECIES guatemalensis

SPECIES gymnotis

SPECIES hooperi

SPECIES hylocetes

SPECIES interparietalis

SPECIES keeni

SPECIES leucopus

SPECIES levipes

SPECIES madrensis

SPECIES maniculatus

SPECIES mayensis

SPECIES megalops

SPECIES mekisturus

SPECIES melanocarpus

SPECIES melanophrys

SPECIES melanotis

SPECIES melanurus

SPECIES merriami

SPECIES mexicanus

SPECIES nasutus

SPECIES ochraventer

SPECIES pectoralis

SPECIES pembertoni

SPECIES perfulvus

SPECIES polionotus

SPECIES polius

SPECIES pseudocrinitus

SPECIES sagax

SPECIES sejugis

SPECIES simulus

SPECIES slevini

SPECIES spicilegus

SPECIES stephani

SPECIES stirtoni

SPECIES truei

SPECIES winkelmanni

SPECIES yucatanicus

SPECIES zarhynchus

GENUS Podomys

SPECIES floridanus

GENUS Reithrodontomys

SUBGENUS Reithrodontomys

SPECIES burti

SPECIES chrysopsis

SPECIES fulvescens

SPECIES hirsutus

SPECIES humulis

SPECIES megalotis

SPECIES montanus

SPECIES raviventris

SPECIES sumichrasti

SPECIES zacatecae

SUBGENUS Aporodon

SPECIES brevirostris

SPECIES creper

SPECIES darienensis

SPECIES gracilis

SPECIES mexicanus

SPECIES microdon

SPECIES paradoxus

SPECIES rodriguezi

SPECIES spectabilis

SPECIES tenuirostris

GENUS Scotinomys

SPECIES teguina

SPECIES xerampelinus

GENUS Xenomys

SPECIES nelsoni

SUBFAMILY Sigmodontinae

GENUS Abrawayaomys

SPECIES ruschii

GENUS Abrothrix

SPECIES andinus

SPECIES hershkovitzi

SPECIES illuteus

SPECIES jelskii

SPECIES lanosus

SPECIES longipilis

SPECIES markhami

SPECIES olivaceus

SPECIES sanborni

GENUS Aepeomys

SPECIES lugens

SPECIES reigi

GENUS Akodon

SPECIES aerosus

SPECIES bogotensis

SUBGENUS Akodon

SPECIES affinis

SPECIES albiventer

SPECIES aliquantulus

SPECIES azarae

SPECIES boliviensis

SPECIES cursor

SPECIES dayi

SPECIES dolores

SPECIES fumeus

SPECIES iniscatus

SPECIES juninensis

SPECIES kofordi

SPECIES leucolimnaeus

SPECIES lindberghi

SPECIES lutescens

SPECIES molinae

SPECIES mollis

SPECIES montensis

SPECIES mystax

SPECIES neocenus

SPECIES oenos

SPECIES orophilus

SPECIES paranaensis

SPECIES pervalens

SPECIES reigi

SPECIES sanctipaulensis

SPECIES serrensis

SPECIES simulator

SPECIES spegazzinii

SPECIES subfuscus

SPECIES surdus

SPECIES sylvanus

SPECIES toba

SPECIES torques

SPECIES varius

SUBGENUS Hypsimys

SPECIES budini

SPECIES siberiae

SUBGENUS Microxus

SPECIES latebricola

SPECIES mimus

GENUS Amphinectomys

SPECIES savamis

GENUS Andalgalomys

SPECIES olrogi

SPECIES pearsoni

SPECIES roigi

GENUS Andinomys

SPECIES edax

GENUS Anotomys

SPECIES leander

GENUS Auliscomys

SPECIES boliviensis

SPECIES pictus

SPECIES sublimis

GENUS Bibimys

SPECIES chacoensis

SPECIES labiosus

SPECIES torresi

GENUS Blarinomys

SPECIES breviceps

GENUS Brucepattersonius

SPECIES albinasus

SPECIES griserufescens

SPECIES guarani

SPECIES igniventris

SPECIES iheringi

SPECIES misionensis

SPECIES paradisus

SPECIES soricinus

GENUS Calomys

SPECIES boliviae

SPECIES callidus

SPECIES callosus

SPECIES expulsus

SPECIES hummelincki

SPECIES laucha

SPECIES lepidus

SPECIES musculinus

SPECIES sorellus

SPECIES tener

SPECIES tocantinsi

SPECIES venustus

GENUS Chelemys

SPECIES delfini

SPECIES macronyx

SPECIES megalonyx

GENUS Chibchanomys

SPECIES orcesi

SPECIES trichotis

GENUS Chilomys

SPECIES instans

GENUS Chinchillula

SPECIES sahamae

GENUS Delomys

SPECIES collinus

SPECIES dorsalis

SPECIES sublineatus

GENUS Deltamys

SPECIES kempi

GENUS Eligmodontia

SPECIES moreni

SPECIES morgani

SPECIES puerulus

SPECIES typus

GENUS Euneomys

SPECIES chinchilloides

SPECIES fossor

SPECIES mordax

SPECIES petersoni

GENUS Galenomys

SPECIES garleppi

GENUS Geoxus

SPECIES valdivianus

GENUS Graomys

SPECIES centralis

SPECIES domorum

SPECIES edithae

SPECIES griseoflavus

GENUS Handleyomys

SPECIES fuscatus

SPECIES intectus

GENUS Holochilus

SPECIES brasiliensis

SPECIES chacarius

SPECIES sciureus

GENUS Ichthyomys

SPECIES hydrobates

SPECIES pittieri

SPECIES stolzmanni

SPECIES tweedii

GENUS Irenomys

SPECIES tarsalis

GENUS Juliomys

SPECIES pictipes

SPECIES rimofrons

GENUS Juscelinomys

SPECIES candango

SPECIES guaporensis

SPECIES huanchacae

GENUS Kunsia

SPECIES fronto

SPECIES tomentosus

GENUS Lenoxus

SPECIES apicalis

GENUS Loxodontomys

SPECIES micropus

SPECIES pikumche

GENUS Lundomys

SPECIES molitor

GENUS Megalomys

SPECIES desmarestii

SPECIES luciae

GENUS Megaoryzomys

SPECIES curioi

GENUS Melanomys

SPECIES caliginosus

SPECIES robustulus

SPECIES zunigae

GENUS Microakodontomys

SPECIES transitorius

GENUS Microryzomys

SPECIES altissimus

SPECIES minutus

GENUS Neacomys

SPECIES dubosti

SPECIES guianae

SPECIES minutus

SPECIES musseri

SPECIES paracou

SPECIES pictus

SPECIES spinosus

SPECIES tenuipes

GENUS Necromys

SPECIES amoenus

SPECIES benefactus

SPECIES lactens

SPECIES lasiurus

SPECIES lenguarum

SPECIES obscurus

SPECIES punctulatus

SPECIES temchuki

SPECIES urichi

GENUS Nectomys

SPECIES apicalis

SPECIES magdalenae

SPECIES palmipes

SPECIES rattus

SPECIES squamipes

GENUS Neotomys

SPECIES ebriosus

GENUS Nesoryzomys

SPECIES darwini

SPECIES fernandinae

SPECIES indefessus

SPECIES swarthi

GENUS Neusticomys

SPECIES monticolus

SPECIES mussoi

SPECIES oyapocki

SPECIES peruviensis

SPECIES venezuelae

GENUS Noronhomys

SPECIES vespuccii

GENUS Notiomys

SPECIES edwardsii

GENUS Oecomys

SPECIES auyantepui

SPECIES bicolor

SPECIES catherinae

SPECIES cleberi

SPECIES concolor

SPECIES flavicans

SPECIES mamorae

SPECIES paricola

SPECIES phaeotis

SPECIES rex

SPECIES roberti

SPECIES rutilus

SPECIES speciosus

SPECIES superans

SPECIES trinitatis

GENUS Oligoryzomys

SPECIES andinus

SPECIES arenalis

SPECIES brendae

SPECIES chacoensis

SPECIES delticola

SPECIES destructor

SPECIES eliurus

SPECIES flavescens

SPECIES fornesi

SPECIES fulvescens

SPECIES griseolus

SPECIES longicaudatus

SPECIES magellanicus

SPECIES microtis

SPECIES nigripes

SPECIES stramineus

SPECIES vegetus

SPECIES victus

GENUS Oryzomys

SPECIES albigularis

SPECIES alfaroi

SPECIES angouya

SPECIES auriventer

SPECIES balneator

SPECIES bolivaris

SPECIES caracolus

SPECIES chapmani

SPECIES couesi

SPECIES curasoae

SPECIES devius

SPECIES dimidiatus

SPECIES emmonsae

SPECIES galapagoensis

SPECIES gorgasi

SPECIES hammondi

SPECIES keaysi

SPECIES lamia

SPECIES laticeps

SPECIES legatus

SPECIES levipes

SPECIES macconnelli

SPECIES maracajuensis

SPECIES marinhus

SPECIES megacephalus

SPECIES melanotis

SPECIES meridensis

SPECIES nelsoni

SPECIES nitidus

SPECIES palustris

SPECIES perenensis

SPECIES polius

SPECIES rhabdops

SPECIES rostratus

SPECIES russatus

SPECIES saturatior

SPECIES scotti

SPECIES seuanezi

SPECIES subflavus

SPECIES talamancae

SPECIES tatei

SPECIES xanthaeolus

SPECIES yunganus

GENUS Oxymycterus

SPECIES akodontius

SPECIES amazonicus

SPECIES angularis

SPECIES caparoae

SPECIES dasytrichus

SPECIES delator

SPECIES hiska

SPECIES hispidus

SPECIES hucucha

SPECIES inca

SPECIES josei

SPECIES nasutus

SPECIES paramensis

SPECIES quaestor

SPECIES roberti

SPECIES rufus

GENUS Paralomys

SPECIES gerbillus

GENUS Pearsonomys

SPECIES annectens

GENUS Phaenomys

SPECIES ferrugineus

GENUS Phyllotis

SPECIES amicus

SPECIES andium

SPECIES bonariensis

SPECIES caprinus

SPECIES darwini

SPECIES definitus

SPECIES haggardi

SPECIES limatus

SPECIES magister

SPECIES osgoodi

SPECIES osilae

SPECIES wolffsohni

SPECIES xanthopygus

GENUS Podoxymys

SPECIES roraimae

GENUS Pseudoryzomys

SPECIES simplex

GENUS Punomys

SPECIES kofordi

SPECIES lemminus

GENUS Reithrodon

SPECIES auritus

SPECIES typicus

GENUS Rhagomys

SPECIES longilingua

SPECIES rufescens

GENUS Rheomys

SPECIES mexicanus

SPECIES raptor

SPECIES thomasi

SPECIES underwoodi

GENUS Rhipidomys

SPECIES austrinus

SPECIES caucensis

SPECIES couesi

SPECIES emiliae

SPECIES fulviventer

SPECIES gardneri

SPECIES latimanus

SPECIES leucodactylus

SPECIES macconnelli

SPECIES macrurus

SPECIES mastacalis

SPECIES modicus

SPECIES nitela

SPECIES ochrogaster

SPECIES venezuelae

SPECIES venustus

SPECIES wetzeli

GENUS Salinomys

SPECIES delicatus

GENUS Scapteromys

SPECIES aquaticus

SPECIES tumidus

GENUS Scolomys

SPECIES melanops

SPECIES ucayalensis

GENUS Sigmodon

SUBGENUS Sigmodon

SPECIES alleni

SPECIES arizonae

SPECIES fulviventer

SPECIES hirsutus

SPECIES hispidus

SPECIES inopinatus

SPECIES leucotis

SPECIES mascotensis

SPECIES ochrognathus

SPECIES peruanus

SPECIES planifrons

SPECIES toltecus

SPECIES zanjonensis

SUBGENUS Sigmomys

SPECIES alstoni

GENUS Sigmodontomys

SPECIES alfari

SPECIES aphrastus

GENUS Tapecomys

SPECIES primus

GENUS Thalpomys

SPECIES cerradensis

SPECIES lasiotis

GENUS Thaptomys

SPECIES nigrita

GENUS Thomasomys

SPECIES apeco

SPECIES aureus

SPECIES baeops

SPECIES bombycinus

SPECIES caudivarius

SPECIES cinereiventer

SPECIES cinereus

SPECIES cinnameus

SPECIES daphne

SPECIES eleusis

SPECIES erro

SPECIES gracilis

SPECIES hudsoni

SPECIES hylophilus

SPECIES incanus

SPECIES ischyrus

SPECIES kalinowskii

SPECIES ladewi

SPECIES laniger

SPECIES macrotis

SPECIES monochromos

SPECIES niveipes

SPECIES notatus

SPECIES onkiro

SPECIES oreas

SPECIES paramorum

SPECIES popayanus

SPECIES praetor

SPECIES pyrrhonotus

SPECIES rhoadsi

SPECIES rosalinda

SPECIES silvestris

SPECIES taczanowskii

SPECIES ucucha

SPECIES vestitus

SPECIES vulcani

GENUS Wiedomys

SPECIES pyrrhorhinos

GENUS Wilfredomys

SPECIES oenax

GENUS Zygodontomys

SPECIES brevicauda

SPECIES brunneus

SUBFAMILY Tylomyinae

GENUS Nyctomys

SPECIES sumichrasti

GENUS Otonyctomys

SPECIES hatti

GENUS Ototylomys

SPECIES phyllotis

GENUS Tylomys

SPECIES bullaris

SPECIES fulviventer

SPECIES mirae

SPECIES nudicaudus

SPECIES panamensis

SPECIES tumbalensis

SPECIES watsoni

Synonyms:


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