Comments: | P. boylii species group. Many taxa previously consigned as subspecies or full synonyms by Osgood (1909) or Hall (1981) have been elevated to species: namely P. attwateri (see Schmidly, 1973), P. aztecus (see Alvarez, 1961; Carleton, 1979; Hooper, 1968), P. baetae (see Bradley and Schmidly, 1987; Schmidly et al., 1988), P. levipes (see Schmidly et al., 1988), P. madrensis (see Carleton, 1977; Carleton et al., 1982), P. simulus (see Carleton, 1977), P. sagax (see Bradley et al., 1996a), and P. spicilegus (see Carleton, 1977). Others have been realigned with other species—cordillerae and evides under P. aztecus (Carleton, 1979); ambiguus under P. levipes (Castro-Campillo et al, 1999; Schmidly et al., 1988); sacarensis under P. baetae (Bradley et al., 2000); and penicillatus under P. nasutus (Diersing, 1976). These changes in status and synonymy of taxa once associated under one species sensu Osgood (1909) involve the systematics of at least three species groups by current understanding. Chromosomal and molecular information suggests that yet other forms in W México await differential characterization as species (see overview by Tiemann-Boege et al., 2000). Karyotypic variation of P. boylii and its kin investigated by Houseal et al. (1987), Lee et al. (1972), and Schmidly and Schroeter (1974); biochemical and molecular variation by Avise et al. (1974), Kilpatrick and Zimmerman (1975), Rennert and Kilpatrick (1986, 1987), Sullivan et al. (1991), Tiemann-Boege et al. (2000); morphological variation by Bradley and Schmidly (1987), Carleton (1977, 1979), and Schmidly et al. (1988). |
Offspring: | Synonyms:
gaurus Elliot, 1903 glasselli Burt, 1932 major (Rhoads, 1893) metallicola Elliot, 1904 parasiticus Elliot, 1904 pinalis (Miller, 1893) robustus (J. A. Allen, 1893) rowleyi (J. A. Allen, 1893) utahensis Durrant, 1946
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