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SPECIES Microtus (See comments.) umbrosus

Author:Merriam, 1898.
Citation:Proc. Biol. Soc. Wash., 12: 107.
Common Name:Zempoaltépec Vole
Type Locality:México, Oaxaca, Mt Zempoaltépec, 8200 ft (2500 m).
Distribution:Cerro Zempoaltépec, 1800-3000 m, NC Oaxaca, México.
Status:IUCN – Lower Risk (lc).
Comments:Type species and sole member of Orthriomys, a taxon typically identified as a subgenus of Microtus (Anderson, 1959; Bailey, 1900; Hall and Cockrum, 1953; R. A. Martin, 1995; Zagorodnyuk, 1990). Nonbanded karyotype reported by Cervantes et al. (1994), who noted its agreement with the complement hypothesized as ancestral for Arvicolinae. Viewed as cladistically basal to other North American pitymyines (subgenera Herpetomys, Pedomys, and Pitymys) based on phylogenetic study of molar traits (R. A. Martin, 1995); more removed from pitymyine species according to mitochrondrial gene sequences and weakly associated with M. chrotorrhinus (Conroy et al., 2001). Conroy et al. (2001) suggested that M. umbrosus represents an independent invasion of southern latitudes, the other consisting of species related to M. pinetorum, subgenus Pitymys (M. guatemalensis, M. oaxacensis, M. quasiater). See Frey and Cervantes (1997, Mammalian Species, 555).
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